There are germs and mold spores floating around in the air all the time everywhere. Even if you washed your hands to perfect sterility, you’re still going to leave some skin cells on the towel along with the water. So now you have a porous towel surface, with water and food (dead skin cells) hanging there getting landed on by mold and bacteria floating around in the air. Perfect growing surface for any that happen to land on the towel, so eventually the towel goes sour, even if you’re hand washing perfects and not adding any organisms via your hands. The sour smell is the pee and poop of bacteria living in it.
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