Air defense at marshaling yards, which were major targets. Also, at bridges. Plain track is easy to fix. Basically, if your enemy has free roam of your skies, then you are screwed logistically. Especially since after major targets are hit, there is no reason not to take potshots at targets of opportunity.
Modernly, it is about keeping enough threat up that they have to make attacks from longer away and be pickier about targets.
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