Engine is aligned to be straight* when the plane is in a cruising attitude.
*Engine is aligned to produce min drag in cruise. If engine thrust is not on center of mass you might change alignment slightly to minimize overall drag from trim to counter torque.
Which plane(s) are you asking about specifically and during which stage of flight. Many people planes have the engine nose up on the ground for example. Which end of the engine are you asking about? That kind of engine?
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