Offense controls what is happening on the field.
They can run plays that target fresh players and give tired players a bit of a break. They can sub players in an out based on how fresh they are.
The defense reacts to the offense. If their defensive backs are winded, the offense isn’t going to call a run play to give them a break. They will keep attacking the secondary. If a defensive player is a little banged up, the offense will target that player.
Sure, the defense can also substitute players, but only to an extent. If the offense comes out with five receivers and is throwing the ball deep, the defense needs to react accordingly.
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