To add to several really good answers:
– malaria and other tropical diseases are definitely a factor in parts of sub-saharan Africa.
– lack of navigable rivers and ports. Africa’s coastline is short compared to its area and the rift valley cannot be navigated by boat. This contrasts with Europe which has many large rivers that criss cross the continent and many inlets and natural ports.
– the Sahara desert. The Egyptian civilization centered around the Nile and was blocked from cultural expansion into the continent by the lack of water away from that river.
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