how did some freshwater fish like sturgeon end up in far-apart places (like the Great Lakes and Caspian Sea, for sturgeon) with no fresh-water path between them?

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how did some freshwater fish like sturgeon end up in far-apart places (like the Great Lakes and Caspian Sea, for sturgeon) with no fresh-water path between them?

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Specifically with sturgeon there are species capable of surviving in saltwater. They are also a very old class of fish, so old that they would have been around just before pangea split up. The likely could have bridged those short saltwater gaps easily. They have speciated since in different regions.

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