The intake valve opens due to the camshaft rotation causing the lob to push down on the spring. When the intake valve opens the piston goes down at the same time drawing in the air due to vacuum. This creates what is know as manifold vacuum. Depending on how far open the throttle valve is will allow a certain amount of air for the “vacuum”, know as mass air flow. Depending on these values the engine control module feeds a signal to the injectors to deliver a certain amount of fuel, this in combination with correct spark timing causes combustion.
Edit: source; automotive tech for 20+ years
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