gravity may have less influence underwater but buoyancy has more. additionallym “up” is where light (and falling food) comes from while “down” is where the ground (and resting food) is so they remain important orientation points. moving between the surface and the depth changes the environment more than moving the same distance sideways
the direction of the light matters because predators tend to sneak up on prey along that axis so their camoflage works. a fish with a pale bottom is harder to see when it’s above you, a fish with a dark top is harder to see when it’s below you
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