Usually its about moving fluids around. While it happens much slower than with water, oils do evaporate and dry out. Most machinery relies on operation to keep fresh oil coating everything. This plays two roles, it maintains lubrication for the next use and keeps a protective film on metals that prevents rust from forming.
Most vehicles are also battery powered. Batteries lose charge over time even when not being used. Turning on the equipment will run the alternator and recharge the battery. This can also be done by just attaching a battery tender to the battery.
Also, quiet, dark, still, and enclosed areas are often viewed by various living things as shelter. They will move in. They will chew up wires. They will crap on everything and jam various nest materials and food for storage area. Damned mice and squirells…..
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