The drivers of the trains know what sections of track require a token. And the track will have signs telling the driver to stop if they don’t have a token.
So if a driver reaches a sign that tells them to stop unless they have a token, then they will stop at the sign. And will then have to figure out where they messed up.
If a driver knows they need to drop-off / pick-up a token at a signal box or platform and there is no one there, then they’ll wait at that point and radio/telephone or just get out and find the member of staff.
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