Lets start longer ago, Citys have been build along Rivers or next to the oceans. Simply because it was far easier to transport stuff this way.
Then with existent Citys Railroads have been build from city to city.
Then since there was a transport option new citys have been build along the railroads.
That was Europe
In America, well i am no historican, but i think that the Railway from East to west coast was necessary to transport the build material. And after the railway have been build it happened the same as in europe, and new Citys along the railroad made sense.
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