This is an active area of research as we do not know what causes it, or even if there is a common reason. The conditions in captivity is very different from the conditions in the wild. The area is smaller, there is less predators and live pray, the animals is watches very often, etc. This can result in very different life expectancy and behavior, in some cases for the better and in some cases for the worse. One factor which have been recently discovered is that some animals show a longer life span when kept as work animals rather then as pets or display animals. But again this only applies to some animals.
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