How do actors in comedies time their lines for laughs without a studio audience?

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How do actors in comedies time their lines for laughs without a studio audience?

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Comedic actors will have had a lot of practice with live audiences in their history as performers and a decent idea of how funny the lines are, and how long of a laugh they’ll get. Also, people laugh out loud less when watching a TV show alone than when in the audience of a comedian.