In living animals, age can be estimated off things like bone density/growth plate fusion, how worn teeth are, in animals like tortoises that have shells, the shells have rings like a tree so can be counted, general size of an animal, and extrapolation from known ages of other animals of similar type. Like if there’s a great white shark that’s 15 feet long and is known to be 30 years old, and was known to grow about 6 inches a year (all numbers hypothetical), and then another shark is seen that’s 25 feet long, an age can be estimated from the average.
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