Some roles are just based on body type. A soldier is never going to become a queen. But others do actually change roles. Ants communicate through chemicals and if an ant in one role goes awhile and doesn’t encounter much of a chemical associated with a role, it can change to that role. So, imagine half the foragers get eaten by an anteater. Other ants slowly notice they’re not encountering enough forager chemicals and might switch roles. That eventually restores the balance.
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