I just want to add to the other great specific answers that these biological don’t “know” anything, as your quotes imply.
You can think of biology as a chemical Rube Goldberg machine, where everything is a consequence of an upstream event, resulting in a domino effect.
So a chemical in the sperm reacts with a chemical in the egg, that causes it to release a chemical that reacts with the outer layer in a way that causes it to harden.
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