Excess CO2/insufficient O2 is just one proposed cause of yawning, and we still don’t know for sure what it does, what causes it, or why it’s such a conserved trait among mammals.
In terms of fatigue for some species, it’s believed to play a role in signaling tiredness to the herd, so everybody can synchronize their sleep schedules together. But again, we don’t know for sure.
It’s thought by some to be a combination of multiple functions which would explain why it’s such a highly conserved trait- it’s versatile.
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