What pain, besides having no oxygen, is a fish feeling when they’re out of the water?

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What pain, besides having no oxygen, is a fish feeling when they’re out of the water?

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They have a protective slime on them that can dry up or be wiped away. Some species don’t do well after this happens. Not sure about pain though. I’ve farmed salmon and never saw one ‘limp’ even with a big bite out of its side.

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