(https://youtu.be/il8bP4ZS4Vc)
Painting starts at around 4min.
The difference materials alone makes it difficult to match perfectly. Often the cel is shiny while the backbrounds are matte paintings. Also for the purpose of consistency and speed, cels are often painted in broad, flat color blocks, which can contrast the more painterly backgrounds.
Furthermore, as someone in the video mentioned, the animated subject is often made to stand out on purpose.
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