We are evolved from tree climbing creatures that used feet like hands while climbing, similar to tree monkeys today.
As our ancestors moved to grasslands and began walking and running upright our toes became part of how we walk and run, and more so how we balance.
I almost lost 4 toes in a motorcycle accident and ended up having surgery to reattach the tendons in them. It took a few years for feeling to return to my toes and it changed my walking and running strides.
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