What are the mechanisms that enable giant tortoises to have exceedingly long natural life spans?

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Is there a genetic component or limited lack of predators?

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Rate/age of living theory. Raymond Pearl based his Age of living theory upon the earlier work of Max Rubner and proposed that the maximum lifespan of an animal was related to its metabolic rate sometimes viewed as the number of heartbeats. However this may only be part of the picture with the role of reactive oxygen species in cell deterioration and eventually cell death also playing a part. – https://youtu.be/ctEIbPI6A4U

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