Most of the top comments seem to miss the main answer:
Vibration and jostling for busses and cars, air pressure and some vibration for planes.
You’ll notice this on land if you go from an old or poorly designed car to one with a more comfortable ride. In the air, if you ride on Boeings Dreamliner, which uses better understanding of jet lag and fatigue to be less exhausting, you’ll see it there.
There are other factors, mentioned by others, such as noise, changing sights, unchanging positions, and more. But jostling and air pressure are the biggest.
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